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In all ordinary cases the glands alone are susceptible to excitement. When excited, they do not themselves move or change form, but transmit a motor impulse to the bending part of their own and adjoining tentacles, and are thus carried towards the centre of the leaf. Stimulants applied to the glands of the short tentacles on the disc indirectly excite movement of the exterior tentacles, for the stimulus of the glands of the disc acts on the bending part of the latter tentacles, near their bases, and does not first travel up the pedicels to the glands, to be then reflected back to the bending place. Some influence, however, does travel up to the glands, causing them to secrete most copiously, and the secretion to become acid, just such an influence as that which in animals is transmitted along the nerves to glands, modifying their power of secretion, independently of the condition of the blood-vessels. Over organic substances that yield soluble matter the tentacles remain clasped for a much longer time than over those not acted upon by the secretion, or over inorganic objects. That they have the power of rendering organic substances soluble, that is, that they have the power of digestion, is no longer a question of dispute. They certainly have this power, acting on albuminous compounds in exactly the same manner as does the gastric juice of mammals, the digested matter being afterwards absorbed. In animals the digestion of albuminous compounds is effected by means of a ferment, pepsin, together with weak hydrochloric acid, though almost any acid will serve, yet neither pepsin nor an acid by itself has any such power. It has been observed that when the glands of the disc are excited by the contact of any object, especially of one containing nitrogeneous matter, the outer tentacles and often the blade become inflected, the leaf thereby becoming converted into a temporary cup or stomach. The discal glands then secrete more copiously, the secretion becoming acid, and, moreover, some influence being transmitted by them to the glands of the exterior tentacles, causing them to emit a more abundant secretion, which also becomes acid. This secretion is to a certain extent antiseptic, as it checks the appearance of mould and infusoria, and in this particular acts like the gastric juice of the higher animals, which is known to arrest putrefaction by destroying the microzymes.

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