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Heterostylous plants may possibly be the means of establishing my view that the law of sexual complements holds good for every kind of living thing. Darwin first, and after him many other investigators have proved that in heterostylous plants fertilisation has the best results, or, indeed, may be possible only when the pollen from a macrostylous flower (a flower with the shortest form of anthers and longest pistil) falls on the stigma of a microstylous blossom (one where the pistil is the shortest possible and the stamens at their greatest length), or vice versâ. In other words, if the best result is to be attained by the cross-fertilisation of a pair of flowers, one flower with a long pistil, and therefore high degree of femaleness, and short stamens must be mated with another possessing a correspondingly short pistil, and so, with the amount of femaleness complementary to the first flower, and with long stamens complementary to the short stamens of the first flower. In the case of flowers where there are three pistil lengths, the best results may be expected when the pollen of one blossom is transmitted to another blossom in which the stigma is the nearest complement of the stigma of the flower from which the pollen came; if another combination is made, either naturally or by artificial fertilisation, then, if a result follows at all, the seedlings are scanty, dwarfed and sometimes infertile, much as when hybrids between species are formed.