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Both of these theories assume that migration is an ingrained habit, but both have been criticized on biological and geological grounds, so neither should be accepted without qualification as definitely accounting for the origin of bird migration. It is apparent, however, that whether the ancestral home of any species was at the northern or southern limits of its present range, or even in some intermediate region, the search for favorable conditions under which to breed in summer and to feed in winter has been the principal factor underlying the origin of migration.

Theory of photoperiodism

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A modern view based on studies of living behavior, suggests also that there is good reason for believing that migration is an annually induced movement. If such be true then the theory of photoperiodism as propounded by some recent investigators should receive some consideration.

This theory holds as its major premise that quantity of light and length of day are the stimulating causes of migration. Its proponents urge that migration is a phenomenon far too regular to be created anew each season merely under stress of circumstances, such as need for food; and that it begins before the necessity for a change in latitude becomes at all pressing. Swallows, nighthawks, shore birds, and others may start their southward movement while the summer food supply in the North is at peak abundance; while robins, bluebirds, and others may leave an abundant food in the South in spring and press toward northern points when food supplies there are almost entirely lacking and when severe cold and storms are likely to cause their wholesale destruction. The regularity of arrival and departure is one of the most impressive features of migration, and since birds travel in almost strict accordance with the calendar, the proponents of the theory ask: "What phenomenon to which we may attribute the stimulating impulse occurs with such precise regularity as the constantly increasing amount of light in spring?"

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