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In their excellent work on the cockroach, Miall and Denny state that “The epipharynx, which is a prominent part in Coleoptera and Diptera, is not recognizable in Orthoptera” (p. 45). We have, however, found it to be always present in this order (Figs. 26, 27).

We are not aware that any modern writers have described or referred to the epipharynx of the mandibulate orders of insects. Although Dr. G. Joseph speaks of finding taste-organs on the palate of almost every order of insects, especially plant-feeding forms, we are unable to find any specific references, his detailed observations being apparently unpublished.

The epipharynx is so intimately associated with the elongated labium of certain Diptera, that, with Dr. Dimmock, we may refer to the double organ as the labrum-epipharynx; and where, as in the lepidopterous Micropteryx semipurpurella, described and figured by Walter, and the Panorpidæ (Panorpa and Boreus), the labrum seems pieced out with a thin, pale membranous fold which appears to be an extension of the epipharynx, building up the dorsal end of the labrum, this term is a convenient one to use.